The flame will visit the Royal Opera House and the Houses of Parliament following the lighting of a ceremonial cauldron in Trafalgar Square.
Torchbearers will also carry it to see performers preparing for this weekend's Notting Hill Carnival and take a trip on the Docklands Light Railway (DLR) as it travels around the capital.
The day will start off with the cauldron being lit in Trafalgar Square by Claire Lomas, who became the first person to complete a marathon in a bionic suit earlier this year.
Ms Lomas, who was left paralysed from the chest down following a horse riding accident in 2007, walked the London Marathon route in a pioneering robotic suit raising ?204,000 for Spinal Research.
She will light the cauldron using the flame that was created at Scafell Pike.
Three other Paralympic flames were struck by Scouts on the summits of the highest peaks of the home nations on Tuesday.
As well as Scafell Pike in England, they were lit on Snowdon in Wales, Ben Nevis in Scotland and Northern Ireland's Slieve Donard.
The flames will rest at Paralympic celebrations in Belfast, Edinburgh and Cardiff along with London before heading to Stoke Mandeville, where they will be united during a special ceremony on August 28 before a 24-hour relay to the opening of the Games at the Olympic Stadium takes place.
As the start of the Games draws nearer, today will also see the Agitos, the official symbol of the 2012 Paralympics, be suspended from Tower Bridge, where it will remain until September 15.
Marking the lighting of the cauldron in Trafalgar Square, Prime Minister David Cameron said he was looking forward to an "incredible" Games.
He said: "After a fortnight of Olympics withdrawal symptoms, it's time to dust off the GB flags and get ready for two more weeks of spectacular sport. The London Paralympic Games 2012 are finally here.
"Already this is looking to be an incredible event. Just like the Olympics, we're going to see international friendship, fierce rivalry and the sheer sporting genius of athletes who have trained day in, day out for years. But we can't deny there's something particularly inspiring about the Paralympics.
"Why? Because being a Paralympian takes an extra measure of guts and steel. A lot of the athletes arriving in London would have been told from a young age about everything they can't do - and they decided to throw everything into what they can do, whether that's smashing a ball over the net, running on prosthetics or swimming faster than the rest.
"That example - of overcoming the difficulties you've been handed, pushing yourself to the limits, going way beyond the expectations others have set for you - is truly inspiring.
"So I want to wish everyone competing in these Games - and particularly ParalympicsGB - the best of luck. The whole country will be cheering you on."
London Mayor Boris Johnson said: "With just five days to go, London is ready to make history and host the most successful Paralympic Games ever seen.
"So get involved, get out on the streets and show your support as the Paralympic Flame, and all the incredible endeavours it represents, makes its way through London."
The day will also see celebrations taking place in Plymouth as the city's flame ambassador, 14-year-old wheelchair athlete Doaa Shayea, and the city's Lord Mayor will take part in a procession after returning by train from the ceremony in Trafalgar Square.
And later the name of the flagbearer for ParalympicsGB during the opening ceremony of the Games will also be announced.
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